ACKNOWLEDGMENTS
This research was supported by the Science Foundation of Jiangsu Province, China 116 [BE2017037], and the Forestry Science and Technology Innovation and Popularization Project in 117 Jiangsu Province, China[LYKJ[2017]14,LYKJ[2018]29, LYKJ[2018]43]. We are grateful to Dr. Sun Zhong-Shuai for his valuable suggestions on the discussion. We are also grateful to Xu Zi-Kun, Shi Cai-Xia, Ke Xian-Shi and Qiu Jing for their help with the field work, and Ren Qing-Hong for sample collection in Korea. Besides, we are thankful to Shang Tao and Cong Rui for help in laboratory work.