Author contributions
XH conducted eDNA metabarcoding, data analysis, and paper writing. SG collected samples, analyzed morpho-taxonomy and DNA barcoding data in the BOLD database, and contributed to manuscript writing. CA, TS, and JP co-conceived the study. CA led the funding proposal, supervised laboratory aspects of the work, and contributed feedback on data analyses. TS led field-planning and the collection of geochemistry data as well as provided critical input to data analyses. MH provided expertise and laboratory assistance related to the DNA barcoding aspects of the project. KM provided expertise and laboratory assistance related to eDNA metabarcoding. KW provided expertise related to statistical analyses. All authors provided feedback on manuscript drafts.