Acknowledgements
This study was supported by supported by the Jinying’s Star Talent Program (Grant no. R2018PY-JX003) from Guangdong Academy of Agricultural Sciences, the Guangdong Key S&T Program (Grant no. 2019B020217002) from Department of Science and Technology of Guangdong Province, the discipline team building projects of Guangdong Academy of Agricultural Sciences in the 13th Five-Year Period.