Author Contributions
F.G. and Y.S. conceived the original screening and research plans; H.G. and Y.S. supervised the experiments; Q.W. and X.L. performed most of the experiments; Q.W., E.Y. conducted the field sampling; Q.W. and H.G. designed the experiments and analyzed the data; Q.W., Y.S. and K.Z-S. wrote the article with contributions of all the authors.