ACKNOWLEDGEMENTS
We thank Louis Hadjioannou, Kim Mortega, Mnqobi Mamba, Maneno Mbilinyi, and Phumlile Simelane for assistance in the field, Alan Howland, Colleen Downs, David Ellers-Smith, Matt and Stephanie McGinn, John and Retha Harding, and Neil Baker and the late Liz Baker for assistance with logistics and Andrea Fulgione for helpful comments on the manuscript. We thank The Kingdom of Eswatini’s Big Game Parks, Ezemvelo KZN Wildlife, Tanzania Wildlife Research Institute (TAWIRI), Tanzania Commission for Science and Technology (COSTECH), Tanzania National Parks (TANAPA), and Mpumalanga Tourism and Parks Agency for research permits. Funding was provided by FP7 Marie Curie Reintegration Grant No. 268316 and a University of Cyprus Research Grant (ANGK), an AG Leventis Foundation grant (MM) and by the AP Leventis Ornithological Research Institute, Jos, Nigeria (ECN and BOO).