Acknowledgements

This study was supported by the National Natural Science Foundation of China (No. 31400361, No. 31901220), the Training Fund of Guangdong Institute of Applied Biological Resources for PhDs, Masters and Postdoctoral Researchers (GIABR-pyjj201703), the GDAS’ Project of Science and Technology Development (2019GDASYL-0105044), the Biodiversity Survey and Assessment Project of the Ministry of Ecology and Environment, China (2019HJ2096001006) and the GDAS’ Special Project of Science and Technology Development (2018GDASCX-0107).