Author Contributions

J. Liang, Z. Zhang and H. Hu conceived and designed the study. J. Liang, Z. Ding, G, Lie and Z. Zhou performed the field survey. J. Liang, Z. Ding and Z. Zhou analyzed the data. J. Liang, Z. Ding and P.B. Singh wrote the paper. All authors reviewed drafts of the paper and agreed on the main conclusions of the manuscript.