Author Contributions
This study represent a revised version of the MSc thesis of LRT. LRT, ET, LM, AB and HK designed the research and LRT, ET, LM and HK conducted fieldwork. LRT, LM, ET, SM and LN did lab work and LRT, LM, SM and SB analysed the data. LRT wrote the manuscript in collaboration with HK and ET and all other co-authors edited and commented.