Author Contributions
SB organized the expeditions and sampling. SB and MS devised the project and supervised the theses of AS and EP. Species identification was performed by SB and JS. The COI data was generated by AS and EP, the ddRAD data by EP, the morphological data by AS and SB; the MALDI-TOF data by EP, SB and SR. The genetic data was analyzed by MS and EP, the MALDI-TOF data by SR and JP. PMA conducted the distribution modelling. MS, EP and SB drafted the first version of the manuscript, all authors contributed significantly to the final version.