Author Contributions
SB organized the expeditions and sampling. SB and MS devised the project
and supervised the theses of AS and EP. Species identification was
performed by SB and JS. The COI data was generated by AS and EP, the
ddRAD data by EP, the morphological data by AS and SB; the MALDI-TOF
data by EP, SB and SR. The genetic data was analyzed by MS and EP, the
MALDI-TOF data by SR and JP. PMA conducted the distribution modelling.
MS, EP and SB drafted the first version of the manuscript, all authors
contributed significantly to the final version.