Author contributions
LJ conceptualized the project and along with KK, IM and AS designed the experiment. AS performed the experiments, analyzed the data, and wrote the manuscript. DJ contributed in RNA-seq bioinformatics pipeline and analysis. APD contributed with the HPLC analysis of anthocyanins. NS performed metabolic profiling with LC-MS. LJ, KK, IM and RVE contributed in the editing and proofreading of the manuscript draft. All authors have read and approved the manuscript.