Author Contributions
Xinmin An designed and managed the project. Rengang Zhang led the genome assembly and downstream analyses. Jianfeng Mao designed and led evolutionary analyses. Steven H. Strauss contributed to scientific analysis and interpretation, and edited manuscript. Stephen R. Keller participed in scientific interpretation and writing the manuscript. Kai Gao and Y.L. created anther plant. Kai Gao, Zhong Chen, J.L., X.Y., X.Y.Y., J.Z., T.Y.Z., T.G., S.H., D.Y.M., W.U.K., B.G., S.W.L., and N.S.M. prepared and collected all plant materials. J.W., B.Q.L., W.H.L., and Q.Z.Y. prepared RNA samples. P.R., M.X.Y., and L.X.J., preformed transcriptome assembly and anlysis. Xinmin An, Kai Gao, and Zhong Chen wrote the manuscript with input from other authors. All authors approved the manuscript before submission.