Author Contributions
J.C.Z and X.W.P. designed the experiments. J.C.Z., Y.F.C., Y.L.W. , Y.L. and T.Q.H. analyzed the data. X.W.P. and J.C.Z. wrote the manuscript. J.C.Z., Y.F.C., Y.L., T.Q.H., Y.L.W., Y.E.Z and X.W.P. revised the manuscript. All authors approved the final version of the manuscript.