Acknowledgements

The authors acknowledge the financial support of the National Natural Science Foundation of China (22038004, 22078069 and 22178069), the Guangdong Basic and Applied Basic Research Foundation (2019B151502038 and 2021A1515012354), Guangdong Provincial Key Research and Development Program (2020B1111380002), and the Guangdong Provincial Key Laboratory of Plant Resources Biorefinery (2021GDKLPRB-K05).