Author Contributions
N.S.-M., T.J.B., J.C.B. and P.M.D. conceived the research and designed the study. S.S., I.A.S.-C., K.L.C., A. Lemma, F.A., A. Lu, T.J.B., J.C.B., and N.S.-M. collected field gelada samples and data, facilitated by A. A. H. P.M.D. performed photo analyses. P.M.D., K.L.C., and N.S.-M. generated genomic data. P.M.D. and N.S.-M. performed genomic analyses. P.M.D., J.C.B., and N.S.-M. wrote the paper. All authors revised and approved the final manuscript.