Authors’ contributions

Yu Wu performed the mostly experiments, analyzed the data, drafted the manuscript. Dai Li and Jian Xie analyzed the data, and critically revised the manuscript. Han Wang, Yan Meng and Guosheng Wu contributed to the data analysis and interpretation. Xiaoming Deng conceived and designed the study. Xiaojian Wan conceived and designed the experiments, analyzed the data, edited the manuscript. All authors substantially contributed to editing, revising, and finalizing the manuscript. All authors read and approved the final manuscript.