Author contributions
JFO, JGS, MR and JR, conceived the study. CCC, JFO, MG and JGS, performed samplings. CCC, JFO performed wet lab work. CCC, JFO and JV performed the bioinformatic analyses. CCC, JFO, MR and JR, interpreted the genomic data. MG, JGS discussed on biological data. CCC, JFO, MR and JR drafted the first version of the manuscript. All authors revised and approved the final version of the manuscript.