References
  1. Amoros, C., & Roux, A. L. (1988). Interaction between water bodies within the floodplains of large rivers: function and development of connectivity. Münstersche Geographische Arbeiten29 (1), 125-130.
  2. Aoyama, K., Yoshida, T., Harada, A., Noguchi, M., Miya, H., & Shibata, H. (2011). Changes in carbon stock following soil scarification of non-wooded stands in Hokkaido, northern Japan. Journal of forest research16 (1), 35-45.
  3. Armstrong, J. B., Schindler, D. E., Ruff, C. P., Brooks, G. T., Bentley, K. E., & Torgersen, C. E. (2013). Diel horizontal migration in streams: juvenile fish exploit spatial heterogeneity in thermal and trophic resources. Ecology, 94 (9), 2066-2075.
  4. Baranyi, C., Hein, T., Holarek, C., Keckeis, S., & Schiemer, F. (2002). Zooplankton biomass and community structure in a Danube River floodplain system: Effects of hydrology. Freshwater Biology, 47 (3), 473–482.
  5. Bellmore, J. R., Baxter, C. V., Martens, K., & Connolly, P. J. (2013). The floodplain food web mosaic: A study of its importance to salmon and steelhead with implications for their recovery.Ecological Applications , 23(1), 189–207.
  6. Borcard, D., Legendre, P., & Drapeau, P. (1992). Partialling out the spatial component of ecological variation. Ecology73 (3), 1045-1055.
  7. Endo, H., Uno, H., Kishida, O., Morita, K. (2023). Habitat characteristics of the Sakhalin taimen Parahucho perryi in the Butokamabetsu River, northern Hokkaido, Japan, Japanese Journal of Conservation Ecology (in Japanese)
  8. Ferrier, S., Watson, G., Pearce, J., & Drielsma, M. (2002). Extended statistical approaches to modelling spatial pattern in biodiversity in northeast New South Wales. I. Species-level modelling. Biodiversity & Conservation11 (12), 2275-2307.
  9. Ferrier, S., Manion, G., Elith, J., & Richardson, K. (2007). Using generalized dissimilarity modelling to analyse and predict patterns of beta diversity in regional biodiversity assessment. Diversity and distributions13 (3), 252-264.
  10. Gallardo, B., Dolédec, S., Paillex, A., Arscott, D. B., Sheldon, F., Zilli, F., … & Comín, F. A. (2014). Response of benthic macroinvertebrates to gradients in hydrological connectivity: a comparison of temperate, subtropical, Mediterranean and semiarid river floodplains. Freshwater Biology59 (3), 630-648.
  11. Gaufin, A.R., Clubb, R., Newell, R. (1974). Studies on the tolerance of aquatic insects to low oxygen concentrations, The Great Basin Naturalist , 34 (1), 45-59.
  12. Gilbert, B., & Lechowicz, M. J. (2004). Neutrality, niches, and dispersal in a temperate forest understory. Proceedings of the National Academy of Sciences101 (20), 7651-7656.
  13. Gounand, I., Harvey, E., Little, C. J., & Altermatt, F. (2018). Meta-ecosystems 2.0: rooting the theory into the field. Trends in Ecology & Evolution33 (1), 36-46.
  14. Hamer, A. J., Czeglédi, I., Gál, B., Sály, P., Szalóky, Z., Preiszner, B., & Erős, T. (2023). Hydrology is a major influence on amphibian abundance in a large European floodplain. Freshwater Biology.68 (8)., 1303-1318.
  15. Holgerson, M. A., Duarte, A., Hayes, M. P., Adams, M. J., Tyson, J. A., Douville, K. A., & Strecker, A. L. (2019). Floodplains provide important amphibian habitat despite multiple ecological threats.Ecosphere , 10(9), e02853.
  16. Jeffres, C. A., Opperman, J. J., & Moyle, P. B. (2008). Ephemeral floodplain habitats provide best growth conditions for juvenile Chinook salmon in a California river. Environmental biology of fishes , 83, 449-458.
  17. Junk, W. J., Bayley, P. B., & Sparks, R. E. (1989). The flood pulse concept in river-floodplain systems. Canadian special publication of fisheries and aquatic sciences106 (1), 110-127.
  18. Kanno, Y., Harris, A. C., Kishida, O., Utsumi, S., & Uno, H. (2022). Complex effects of body length and condition on within‐tributary movement and emigration in stream salmonids. Ecology of Freshwater Fish , 31(2), 317-329.
  19. Kawai, T., & Tanida, K. (2005). Aquatic insects of Japan: Manual with keys and illustrations. Second edition. Tokai University Press, Japan . (in Japanese)
  20. Kishida, O., Costa, Z., Tezuka, A., & Michimae, H. (2014). Inducible offences affect predator–prey interactions and life‐history plasticity in both predators and prey. Journal of Animal Ecology , 83(4), 899-906.
  21. Legendre, P., & Legendre, L. (1998). Numerical ecology (2nd English ed.) Elsevier. Amsterdam, The Netherlands .
  22. Leibold, M. A., Holyoak, M., Mouquet, N., Amarasekare, P., Chase, J. M., Hoopes, M. F., … & Gonzalez, A. (2004). The metacommunity concept: a framework for multi‐scale community ecology. Ecology letters7 (7), 601-613.
  23. Levin, S. A. (1992). The problem of pattern and scale in ecology: the Robert H. MacArthur award lecture. Ecology73 (6), 1943-1967.
  24. Manion, G., Lisk, M., Ferrier, S., Lugilde, K. M., Fitzpatrick, M. C., Fitzpatrick, M. M. C., … & Rcpp, I. (2017). Package ‘gdm’. A toolkit with functions to fit, plot, and summarize Generalized Dissimilarity Models: CRAN Repository .
  25. Morand, A., & Joly, P. (1995). Habitat variability and space utilization by the amphibian communities of the French Upper-Rhone floodplain. In Space partition within aquatic ecosystems  (pp. 249-257). Springer, Dordrecht.
  26. Ogawa, A., Shibata, H., Suzuki, K., Mitchell, M. J., & Ikegami, Y. (2006). Relationship of topography to surface water chemistry with particular focus on nitrogen and organic carbon solutes within a forested watershed in Hokkaido, Japan. Hydrological Processes: An International Journal20 (2), 251-265.
  27. Oksanen, J., Blanchet, F. G., Kindt, R., Legendre, P., Minchin, P. R., O’hara, R. B., … & Oksanen, M. J. (2013). Package ‘vegan’. Community ecology package, version2 (9), 1-295.
  28. Peterson, D. W., & Reich, P. B. (2008). Fire frequency and tree canopy structure influence plant species diversity in a forest-grassland ecotone. Plant Ecology194 (1), 5-16.
  29. Power, M. E., Parker, M. S., & Dietrich, W. E. (2008). Seasonal reassembly of a river food web: floods, droughts, and impacts of fish.Ecological Monographs , 78(2), 263-282.
  30. Richardson, J. S. (1991). Seasonal food limitation of detritivores in a montane stream: an experimental test. Ecology, 72(3), 873-887.
  31. Risser, P. G. (1995). The status of the science examining ecotones.BioScience , 45(5), 318-325.
  32. Shibata, H., Sugawara, O., Toyoshima, H., Wondzell, S. M., Nakamura, F., Kasahara, T., … & Sasa, K. (2004). Nitrogen dynamics in the hyporheic zone of a forested stream during a small storm, Hokkaido, Japan. Biogeochemistry69 (1), 83-104.
  33. Smith, T. B., Wayne, R. K., Girman, D. J., & Bruford, M. W. (1997). A role for ecotones in generating rainforest biodiversity.Science, 276(5320), 1855-1857.
  34. Thompson, R., & Townsend, C. (2006). A truce with neutral theory: local deterministic factors, species traits and dispersal limitation together determine patterns of diversity in stream invertebrates. Journal of Animal Ecology75 (2), 476-484.
  35. Tockner, K., Malard, F., & Ward, J. V. (2000). An extension of the flood pulse concept. Hydrological processes14 (16‐17), 2861-2883.
  36. Uno, H., Yokoi, M., Fukushima, K., Kanno, Y., Kishida, O., Mamiya, W., … & Utsumi, S. (2022). Spatially variable hydrological and biological processes shape diverse post‐flood aquatic communities. Freshwater Biology67 (3), 549-563.
  37. Van den Brink, F. W. B., Van Katwijk, M. M., & Van der Velde, G. (1994). Impact of hydrology on phyto-and zooplankton community composition in floodplain lakes along the Lower Rhine and Meuse. Journal of Plankton Research16 (4), 351-373.
  38. van der Maarel, E. (1990). Ecotones and ecoclines are different. Journal of Vegetation Science , 135-138.
  39. Whiles, M. R., & Goldowitz, B. S. (2001). Hydrologic influences on insect emergence production from central Platte River wetlands.Ecological Applications , 11(6), 1829-1842.
  40. Yang, L. H. (2008). Pulses of dead periodical cicadas increase herbivory of American bellflowers. Ecology , 89(6), 1497-1502.
Table 1 SIMPER analysis showing the top ten contributing taxa to the difference between connectivity categories for each season.