ACKNOWLEDGEMENTS
We appreciate the National Plant Specimen Resource Center (NPSRC) and Plant Photo Bank of China (PPBC) for providing occurence data of Lamiaceae species. We are grateful to Dr. Yang Niu for providing photo of Elsholtzia eriostachya. This study was supprted by the Science and Technology Innovation Project of China Southern Power Grid Co., Ltd. (CGYKJXM20220305), Natural Science Foundation of Yunnan Province (202401AT070011), and the Scientific and Technological Innovation Project of Yunnan Forestry Technological College (KY(BS)202301).