In contrast, arthropod groups such as Carabidae and Lycosidae, which showed no clear relationship with the presence of dry refuges, consist mostly of larger, cursorial species. It is possible that these groups are less able to climb the hummocks, instead using their higher mobility to evade flooding and recolonise (Bates, et al. 2006, Lafage, Sibelle, Secondi, Canard and Petillon 2015). As a result, species within Carabidae and Lycosidae that cannot survive flooding are likely filtered out of the community. Meanwhile, flood sensitive species within Linyphiidae and Staphylinidae may persist by moving to the hummocks or other dry refuges. Consequently, we would expect that the Linyphiidae and Staphylinidae contain a higher proportion of species that do not necessarily have wetlands as their main and only habitat. Our data also showed that Carabidae (93%) and Lycosidae (86%) had a much higher proportion of wetland specialists than Linyphiidae (54%) and Staphylinidae (47%). Furthermore, half of the genera classified as habitat generalists responded positively to dry refuges (Amischa,Rybaxis and Savignia but not Bathyphantes, Erichsonius and Oedothorax), whereas none of the genera among wetland specialists showed a positive response to dry refuges despite the much higher number of species. Notice that these genera are primarily represented by a single species each in the data set (Appendix 1).