ACKNOWLEDGEMENTS
We thank Lichao Wang, Qianwen Chu, Fuqi Liu and Minmin Sun for their help with plant cultivation and harvest. We also thank Mingxin Pan, Shengchao Ma and Huaisong Wang for their help with data analyses. This study was supported by the National Natural Science Foundation of China (No. 32171645, 32171551), the Department of Science and Technology of Jilin Province (No. 20240601073RC, 20220101292JC), and the Education Department of Jilin Province (No. JJKH20221171KJ). We thank the anonymous reviewers for their helpful comments on the revision of this manuscript.